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of plant litter and is now being considered as an important driver of decomposition, although uncertainty exists in quantifying its regional and global biogeochemical significance. Changes in the decomposition of plant litter from exposure to UV-B and also UV-A 15–400 nm and visible radiation have potential consequences for the cycling and storage of carbon and other nutrients. As spring comes to Antarctica, the PSC’s melt in the stratosphere and release all of the halogenated compounds that were previously absorbed to the cloud. In the antarctic summer, high energy photons are able to photolyze the halogenated compounds, freeing halogen radicals that then catalytically destroy O. Because Antarctica is constantly surrounded by a polar vortex, radical halogens are not able to be diluted over the entire globe. The ozone hole develops as result of this process. Stratospheric ozone is constantly produced by the action of the sun’s ultraviolet radiation on oxygen molecules . Although ozone is created primarily at tropical latitudes, large-scale air circulation patterns in the lower stratosphere move ozone toward the poles, where its concentration builds up. Schematic illustration of the evolution of ground-level ozone in urban air and its outflow, as a function of distance from emission sources. Dashed curve gives reference using current UV radiation, while the solid curve is for decreased UV radiation expected upon recovery of stratospheric ozone. The green hatched area shows the resultant de-crease in urban ozone, while the red hatched area shows the increase in regional ozone. Smoke from drought-related wildfires and biomass burning as well as increased input of UV-absorbing DOM may reduce underwater exposure to UV radiation see sections 1 and 2, leading to a shallower depth distribution of zooplankton in highly transparent lakes.195 These observations are consistent with the proposition that transparency regulates the relative importance of UV radiation vs. visual predation risk as factors driving vertical migration of zooplankton in lakes.29 Hence, even though UV radiation is harmful to zooplankton and other invertebrates if exposure is high, UV radiation is also a natural environmental cue affecting orientation and vertical distribution in clear-water systems. In addition to changes in quality of water, the critical importance of zooplankton in aquatic food webs means that these changes in behaviour in response to UV radiation have important implications for fishery productivity. Responses to UV radiation alter the depth distribution of zooplankton and thus their vertical overlap in the water column with their algal food resources as well as fish predators. In coastal marine systems, higher quality food with more nutrients can increase tolerance of amphipods shrimp-like crustaceans to UV radiation.22 It has also been demonstrated that amphipods obtain UV-absorbing compounds from their macroalgae food, when exposed to UV radiation.2 Reductions

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