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the large marine predatory fish; air-respiratory would have been obligatory as a result of these waters would have been short of oxygen, since heat water holds less dissolved oxygen than cooler marine water and for the reason that decomposition of vegetation would have used a number of the oxygen. However, in 1987, practically complete fossils of Acanthostega from about 363 Ma confirmed that this Late Devonian transitional animal had legs and both lungs

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